How does wild tomatoes flower?

[imgrimmer]

some days ago I took a closer look on the flowers of borage / starflower also it is a different botanic family the flowers look very similar to tomatoes. (stigma hidden behind the anthers) I read that the stigma only is exposed after the release of pollen by bumblebees, to avoid self pollination.
All of the so called wild species I have grown had hidden stigmas like a normal tomato, I just had the idea that
this is similar in wild tomato species?
It is just a speculation, but maybe exposed stigmas is a new trait in cultivated tomatoes and not the original way of flowering in tomatoes?
I guess someone might have more knowledge on this than me.
thank you

[DarJones]

Wild species use either exerted stigmas or self-incompatibility or both to increase pollination from unrelated plants. The self-incompatible gene is so specific to preventing self pollination that it almost never allows self set of fruit. S. Habrochaites is an example of a species that most often has the S gene, but there are a few lines that are self-compatible. LA2175 is an example of a self compatible S. Habrochaites.

Exerted stigma is most often associated with lines that do not contain the S gene. S. Pimpinellifolium is the direct progenitor of domestic tomato and mostly has exerted stigmas. It is extremely easy to cross S. Pimpinellifolium to domestic tomato and works in either direction. The one trait you almost always get in the segregating offspring is exerted stigmas. The consensus is that domestic tomato was deliberately selected for enclosed stigma because the trait results in larger and better flavored fruits.

[imgrimmer]

Thanks for the explanation! Since borage is from a complete other botanical family, it makes much more sense this way. I guess I will grow some pimpinellifolium for this and other reason